I'm not getting how humidity can prevent or cause pyramiding. Those studies always perplexed me.
If you look at the pictures it's the bones that pyramid, not the scutes. Humidity only helps the scute material from sticking together at the edges and causing lopsided growth of the scutes, but not affecting the bone underneath.
Bones need 4 things to grow, calcium, phosphorus, vitamin D3 and enough calories to untilize the first 3. From what I've always seen it's a deficiency or lack of 1, 2, 3 or all 4 that causes pyramiding.
Glad I could help by posting these pictures
Danny